Lucy Bailey is not a girl to take no for an answer. When she asks her friend Billy Wellington to help her rescue a stray dog, she has no idea of the potential repercussions. A serious crime is committed while Billy is absent from the children's home where he lives and, when suspicion falls on him, the police decide that the safest thing for everybody is to lock him away in a mental institution. Lucy refuses to believe that Billy has done anything wrong, and enlists her cool-headed teenage brother Arnold to help.    

Billie Challinor's mother dies during an air raid, but the child grows up confident that in her jazz musician father, Chas, she has the best dad in the world. They seek refuge from the London Blitz by moving to Leeds, where kindly landlady Liz Morris befriends them: the scarred, wisecracking man who isn't afraid to overstep the mark if the cause is a good one and his clever and resilient little girl.

Eleven-year-old Billy Clegg and his big sister Peggy disappear from their home in Leeds after a suspicious fire in a local mill. The owner says the children caused it by lighting fireworks on Mischief Night. Their widowed mother, Betty, badly injured in the incident, doesn't know if her children are alive or dead. She prefers to think of them as being elsewhere. 

In the summer of 1913, the world seems full of possibility for four very different young men. Young Jean-Baptiste dreams of the day he'll leave his Picardy home and row down-river to the sea. Earnest and hard-working Frank has come to London to take up an apprenticeship in Regent Street. His ambitions are self-improvement, a wife and, above all, a bicycle. Organ scholar Benedict is anxious yet enthralled by the sensations of his synaesthesia.

Warwickshire, 1875 Emerald Pritchard has lived a privileged life with her parents and her younger sister, Abigail, in the stately Astley House on the outskirts of Nuneaton. But all that suddenly changes when her father disappears leaving the family in debt. They are forced to throw themselves on the mercy of Emerald's uncle who begrudgingly allows them to live in a small cottage within the grounds of his farm. 

All Mary wants to do is protect her family. But how can she, when their father keeps tearing it apart? Grieving for their mother and abandoned by their father, Mary and her siblings seek refuge at the foundling school. But the school has its own struggles, short-staffed since Hilary married. When the nuns hire Mary's father, and he arrives with the pregnant 16-year-old he left their mother for, Mary's newfound world is shaken. Can she make a new life for her siblings when their father is set on disrupting it? 

Left on the steps of an orphanage when she was just days old, Nancy Sunday was brought up in hardship—until the kindly Rosalind Carey took her in. Now 18 years old, Nancy is an adopted member of the Carey family. But she can’t help wondering who her parents really were. When Nancy is sent away to finishing school, she finds herself in the midst of London society. There she meets Freddie Ashton—kind and warm-hearted, he might just be the man of Nancy’s dreams. But she knows his wealthy parents would never let him marry a penniless foundling.

Charlotte Booth loves her father and the home they share, which is set high up in the limestone escarpments of Crummockdale. But when a new businessman, Joseph Dawson, enters their lives, both Charlotte and her father decide he's the man for her, and within six months Charlotte marries the dashing mill owner from Accrington. Then a young mill worker is found dead in the swollen River Ribble. With Joseph's business nearly bankrupt, it becomes apparent that all is not as it seems, and Joseph is not the man he pretends to be.

Yorkshire Dales, 1850. As a terrible storm rages, Annabelle Wallis is shocked to find a distressed young woman at her cottage door, heavy with child. Moments later a baby girl is born. But by dawn, the mother has vanished, leaving behind the helpless child wrapped only in a silk peacock shawl. When news spreads that Lady Eliza Hartley, sister to a wealthy estate owner, has been found dead, Annabelle realises the terrible secret she has stumbled on. Terrified she’ll be blamed for Eliza’s death, Annabelle flees to the slums of York, where she plans to raise the orphan as her own.

Julie Monday enjoys a rare moment of happiness when at the age of eight she visits the Essex seaside and meets eleven-year-old Harry Walker. Ten years later, they meet again. Fated to be together, they marry just before the outbreak of the Second World War. But it’s now 1939, Harry has enlisted, and Julie must face the blitz alone with their son. Travelling back from work one day, Julie is caught in the chaos of a direct hit.

Summer, 1936. Newly trained midwife Hazel Price returns to the Yorkshire streets of her childhood, only to find that her modern methods and 'stuck-up' ways bring her into conflict with her family and other formidable residents of Raglan Road. Determined, Hazel battles on, assisting with home deliveries and supporting the local GP. The days are long and hard, but Hazel brings knowledge and compassion to the work she loves.

Kay Duffield's fiancé is about to leave the country, and her own duty with the WAAF is imminent when she becomes a bride. The precious few days she spends with her new husband are quickly forgotten once she starts work as a balloon operator. There she makes friends with shy Emily Bevan, who has left her parents' hill farm in Wales for the first time; down-to-earth Biddy Bachelor, fresh from the horrors of the Liverpool bombing; and spirited Jo Stewart, the rebel among them, whose disregard for authority looks set to land them all in trouble. The girls stay friends through good times and bad, through romantic encounters and, ultimately, tragedy.

Hull, 1940. Widowed Edie Copeland has just arrived on Exchange Street, heavily pregnant and with heavy suitcase in tow. She left York after tragedy struck to make a fresh start, but she's a stranger to this street, and her new landlord doesn't hesitate to remind her of this, widow or not. Luckily, the neighbours are a little more welcoming and Edie is soon made to feel at home by the Maguires and the Scuttles. As air raids sound and the war feels closer than ever, the community has to stick together. But Edie is hiding something, and she doesn't know how much longer her secret is safe.

Summer, 1939. The de Lacey family believe they are ready for the changes the war will bring. Elizabeth, the eldest daughter, is due to return home from her grandparents’ farm in France and is expected to marry Captain Max Coburn. But when Grandpère suffers a stroke, her future is changed. Instead of returning home, Elizabeth chooses to stay and help her grandparents. Max is also stationed in France – but will the war threaten to separate them for ever? Meanwhile, in the village of Nayton, Lucy Storey dutifully cares for her father the stationmaster.

Plymouth, 1941. As sirens blare all around, the Shawbrook family take refuge in a packed shelter. Bombs have already begun to fall through the night sky when they realise their infant son, Freddie, has been forgotten in the rush, left to sleep in his crib. Terrified, Vera, his young mother, races to find him and bring him to safety. The next morning, air raid warden David Shawbrook returns from his watch to find the shelter pulverised, and his family seemingly all dead. Dirty footprints inside their home betray the looters who have rifled through the house.